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The series follows Meredith Grey (Ellen Pompeo), the daughter of an esteemed general surgeon named Ellis Grey, following her acceptance into the residency program at the fictional Seattle Grace Hospital. During her time as a resident, Grey works alongside fellow doctors Cristina Yang (Sandra Oh), Izzie Stevens (Katherine Heigl), Alex Karev (Justin Chambers), and George O'Malley (T. Knight), who each struggle to balance their personal lives with the hectic work and training schedules.

They are overseen during their internship by Miranda Bailey (Chandra Wilson), a senior resident who works for attending Derek Shepherd (Patrick Dempsey), the head of neurosurgery and Meredith's love interest; Preston Burke (Isaiah Washington), the head of cardio who becomes Yang's fiancee; and Richard Webber (James Pickens, Jr. ), the Chief of Surgery and attending general surgeon, and lover of Ellis Grey decades ago. The residents are later joined by Jackson Avery (Jesse Williams) and April Kepner (Sarah Drew), former Mercy-West residents who join Seattle Grace following an administrative merger in the sixth season.

Throughout the first six seasons, Burke, O'Malley and Stevens all depart the series.